COLUMBUS, Ohio (AP) — Trustees of the Columbus Zoo and Aquarium said Friday that financial abuses by four former executives cost the institution more than $630,000.

A forensic analysis ordered by the zoo’s board of trustees showed money from a zoo levy that pays for animal care was not involved.

The review was ordered after a Columbus Dispatch investigation showed apparent improprieties by the zoo’s then-chief executive and financial officers, including misuse of zoo properties and sports tickets.

The review ordered by trustees found a marketing vice president and purchasing director also were involved.

Trustees have pledged to take immediate action to restore public trust.